The stake columns display the back and lay stakes that will be used when placing a back or lay bet on a given selection.

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You can adjust the default value to be displayed in the grid in preferences.

The new default value will be displayed when you next select a market.

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You can quickly adjust the stakes in the grid by overtyping them in the grid or by clicking on one of the preset stake buttons as shown in the image below.

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Using one of the preset stake buttons adjusts the stake in the grid against all selections.

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Double click on a stake button to change its default value.